Sorry, yes, it should crawl the links - they used to do that.
But you can prove it to yourself, by doing what I said - and then report back.
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Posts made by loopyal
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RE: Do search engines crawl links on 404 pages?
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RE: Do search engines crawl links on 404 pages?
Yes and you can test it by creating a page that is linked from nowhere else and then check your logs or analytics